Environmental heterogenity

Environmental heterogenity

fine-grained

a single individual is exposed to several environments, and this exposure always occurs as a temporal sequence. The fitness impact of an environmental state must be a function not only of the genotype and that state, but also of the past environments to which the individual has been exposed and their fitness impacts.

coarse-grained

Fitness is a function of the genotype and the realized environmental state in which the genotype is expressed.